Variations of the tilted guns, as shown on the BE2c above, were also used on the Sopwith 1-1/2 Strutter single seat night fighters (twin Lewis) and the (Hagar, help me on this) "Comic Camel" night fighter (which I believe did NOT have the "hump".

Captain F.W. Honnett, Flight Commander of "A" Flight No. 78 Sqn (HD) RFC, suggested a modification of one of the 1½ Strutters by moving the pilot's seat and all the controls into the observer's position, his argument being poor visibility from the regular pilot's seat. The original pilot's position was faired over, and the plane was equipped with a night searchlight.
The first three 1½ Strutters modified to the new standard by the Southern Aircraft Repair Depot joined 78 Sqn in September 1917. During the night raid over London on the night of October 31st/November 1st 1917 they opposed twenty-two enemy Gothas. 78 Sqn pilots dubbed this unusual plane the 'Comic fighter'. Initially the armament of this aircraft consisted of only a single course Vickers gun; later Comics were equipped with a Lewis gun on a flexible Foster mounting. It should be also mentioned that at least one aircraft, namely B762, had two Lewises on a special fixed mounting and could fire at a 70° angle.
1½ Strutter Comics were intensively used by 78 Sqn until February 1918, flying night intercept missions against Gothas and Giant R-planes. Due to the poor performance of this type, it was never put into series production. At the beginning of 1918 the night fighter version of the famous Sopwith Camel (which ironically received the official name Sopwith Comic) replaced the 1½ Strutter Comic and other obsolete night-fighters in many Home Defence units.

I remember seeing something and I can't remember where...but...there seemed to be two large wheels in the pilot house (gondola) of the airship...one facing forward with an operator and one facing to port with an operator.
The one facing forward (they were like ships wheels) was for the rudder operator and the one facing to port was for the elevator operator.
There was also a throttle man (engineer station), radio station and navigators station.
I do not recall seeing a seat for the Captain and I think the area in the gondola was to small for one.
I will try to find that picture.
The older submarines I think operated with two separate wheels.

I believe that the Goodyear "blimps" still use separate elevator wheels, which are recovered from blimp to blimp as they are retired and a new one is placed in service.  Current elevator wheels date back to the patrol blimps of the '40s.